PNAS

Proceedings of the National Academy of Sciences Template

Write in a clean editor, then format for Proceedings of the National Academy of Sciences in one click — DocuGuru applies the official PNAS template with numbered references and exports a submission-ready PDF plus the editable LaTeX source. Free to start.

About the Proceedings of the National Academy of Sciences format

Proceedings of the National Academy of Sciences is a peer-reviewed journal published by PNAS, covering RNA and protein synthesis mechanisms, Bacterial Genetics and Biotechnology, Neuroscience and Neuropharmacology Research.

PublisherPNAS
Reference styleNumbered (PNAS)
Numbered — [1], [2] in the text
1. A. Smith, B. Jones, C. Lee, A representative article title. Proceedings of the National Academy of Sciences 12, 45–58 (2023).

Formats any DOI in Proceedings of the National Academy of Sciences style. No sign-up.

Publishes research inRNA and protein synthesis mechanisms Bacterial Genetics and Biotechnology Neuroscience and Neuropharmacology Research Monoclonal and Polyclonal Antibodies Research RNA Research and Splicing
ISSN0027-8424
Citation impact (2-yr)8.05
h-index1173
i10-index144,911
Total citations22,890,303
Top institutions publishing hereHarvard University
Journal websitewww.pnas.org
You getA submission-ready PDF and the editable LaTeX source — ready to submit.

Papers published in Proceedings of the National Academy of Sciences per year

4.3K
2014
4.1K
2015
3.9K
2016
4K
2017
4K
2018
4K
2019
4.4K
2020
4.2K
2021
3.9K
2022
3.6K
2023
4K
2024
4.5K
2025

Citation impact of Proceedings of the National Academy of Sciences by publication year

569.2K
2014
497.8K
2015
446.2K
2016
418K
2017
392.5K
2018
345.9K
2019
330.1K
2020
235K
2021
149.5K
2022
99.6K
2023
61.6K
2024
21.3K
2025

Citations each year’s papers have accumulated so far — the most recent years are still building up.

Most-cited papers in Proceedings of the National Academy of Sciences

DNA sequencing with chain-terminating inhibitors

Frederick Sanger, S. Nicklen, Alan Coulson · 1 Dec 1977

A new method for determining nucleotide sequences in DNA is described. It is similar to the "plus and minus" method [Sanger, F. & Coulson, A. R. (1975) J. Mol. Biol. 94, 441-448] but makes use of the 2',3'-dideoxy and arabinonucleoside analogues of the normal deoxynucleoside triphosphates, which act as specific chain-terminating inhibitors of DNA polymerase.…

69,331 citations Cite SaveGo to paper →
Gene set enrichment analysis: A knowledge-based approach for interpreting genome-wide expression profiles

Aravind Subramanian, Pablo Tamayo, Vamsi K. Mootha et al. · 30 Sep 2005

Although genomewide RNA expression analysis has become a routine tool in biomedical research, extracting biological insight from such information remains a major challenge. Here, we describe a powerful analytical method called Gene Set Enrichment Analysis (GSEA) for interpreting gene expression data. The method derives its power by focusing on gene sets, that is, groups of…

57,250 citations Cite SaveGo to paper →
Electrophoretic transfer of proteins from polyacrylamide gels to nitrocellulose sheets: procedure and some applications.

Harry Towbin, T. Staehelin, J. Gordon · 1 Sep 1979

A method has been devised for the electrophoretic transfer of proteins from polyacrylamide gels to nitrocellulose sheets. The method results in quantitative transfer of ribosomal proteins from gels containing urea. For sodium dodecyl sulfate gels, the original band pattern was obtained with no loss of resolution, but the transfer was not quantitative. The method allows…

53,861 citations Cite SaveGo to paper →
Neural networks and physical systems with emergent collective computational abilities.

J. J. Hopfield · 1 Apr 1982

Computational properties of use of biological organisms or to the construction of computers can emerge as collective properties of systems having a large number of simple equivalent components (or neurons). The physical meaning of content-addressable memory is described by an appropriate phase space flow of the state of a system. A model of such a…

19,403 citations Cite SaveGo to paper →
Cluster analysis and display of genome-wide expression patterns

Michael B. Eisen, Paul T. Spellman, Patrick O. Brown et al. · 8 Dec 1998

A system of cluster analysis for genome-wide expression data from DNA microarray hybridization is described that uses standard statistical algorithms to arrange genes according to similarity in pattern of gene expression. The output is displayed graphically, conveying the clustering and the underlying expression data simultaneously in a form intuitive for biologists. We have found in…

16,441 citations Cite SaveGo to paper →

Proceedings of the National Academy of Sciences template — frequently asked questions

How do I write a paper in the Proceedings of the National Academy of Sciences format?
In DocuGuru you write your manuscript in a normal editor — no LaTeX setup required — and select the Proceedings of the National Academy of Sciences template. When you export, DocuGuru compiles the paper into the official PNAS format and hands you a submission-ready PDF along with the editable LaTeX source.
What reference style does Proceedings of the National Academy of Sciences use?
Proceedings of the National Academy of Sciences uses Numbered (PNAS) references, shown as numbered [1], [2] markers in the text. DocuGuru formats every in-text citation and the reference list in this exact style automatically. A reference appears like this: 1. A. Smith, B. Jones, C. Lee, A representative article title. Proceedings of the National Academy of Sciences 12, 45–58 (2023).
Do I need to know LaTeX to submit to Proceedings of the National Academy of Sciences?
No. DocuGuru generates the pnas-new LaTeX class and compiles the PDF for you in the background, so you get a PNAS-ready Proceedings of the National Academy of Sciences document without writing any LaTeX. If you do want it, the LaTeX source is included in the export.
Can I import an existing draft into the Proceedings of the National Academy of Sciences template?
Yes. Paste or upload your current manuscript — Word, LaTeX, Markdown, or plain text — and DocuGuru reflows it into the Proceedings of the National Academy of Sciences format with correct headings, figures, tables, and numbered citations.
Who publishes Proceedings of the National Academy of Sciences?
Proceedings of the National Academy of Sciences is a multidisciplinary journal published by PNAS. DocuGuru's Proceedings of the National Academy of Sciences template matches PNAS's official submission format.
Can I export a submission-ready Proceedings of the National Academy of Sciences PDF?
Yes — DocuGuru produces a PDF built with the official Proceedings of the National Academy of Sciences template (the pnas-new class) that is ready to submit to PNAS, together with the matching LaTeX source files.
How much does the Proceedings of the National Academy of Sciences template cost?
You can start writing in the Proceedings of the National Academy of Sciences template for free. Exporting the final submission-ready Proceedings of the National Academy of Sciences PDF and LaTeX source is part of DocuGuru's paid plans — see the app for current pricing.
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