BMC (BioMed Central)

Zoological Letters Template

Write in a clean editor, then format for Zoological Letters in one click — DocuGuru applies the official BMC (BioMed Central) template with numbered references and exports a submission-ready PDF plus the editable LaTeX source. Free to start.

About the Zoological Letters format

Zoological Letters is a peer-reviewed journal published by BMC (BioMed Central), covering Developmental Biology and Gene Regulation, Neurobiology and Insect Physiology Research, Insect and Arachnid Ecology and Behavior. Over its lifetime it has published 265 papers, which have received 5,395 citations.

PublisherBMC (BioMed Central)
Reference styleNumbered (Vancouver)
Numbered — [1], [2] in the text
1. Smith A, Jones B, Lee C. A representative article title. Zoological Letters. 2023;12(3):45–58.

Formats any DOI in Zoological Letters style. No sign-up.

Publishes research inDevelopmental Biology and Gene Regulation Neurobiology and Insect Physiology Research Insect and Arachnid Ecology and Behavior Marine Ecology and Invasive Species Animal Behavior and Reproduction
ISSN2056-306X
Citation impact (2-yr)2.32
h-index38
i10-index156
Article processing charge$1,680
Open accessYes — indexed in DOAJ
Top institutions publishing hereThe University of Tokyo
Journal websitelink.springer.com
You getA submission-ready PDF and the editable LaTeX source — ready to submit.

Papers published in Zoological Letters per year

1
1990
35
2015
25
2016
22
2017
32
2018
37
2019
16
2020
16
2021
14
2022
23
2023
23
2024
10
2025

Citation impact of Zoological Letters by publication year

0
1990
1.3K
2015
796
2016
558
2017
803
2018
917
2019
285
2020
229
2021
157
2022
208
2023
88
2024
11
2025

Citations each year’s papers have accumulated so far — the most recent years are still building up.

Most-cited papers in Zoological Letters

Bivalve-specific gene expansion in the pearl oyster genome: implications of adaptation to a sessile lifestyle

Takeshi Takeuchi, Ryo Koyanagi, Fuki Gyoja et al. · 18 Feb 2016

INTRODUCTION: Bivalve molluscs have flourished in marine environments, and many species constitute important aquatic resources. Recently, whole genome sequences from two bivalves, the pearl oyster, Pinctada fucata, and the Pacific oyster, Crassostrea gigas, have been decoded, making it possible to compare genomic sequences among molluscs, and to explore general and lineage-specific genetic features and trends…

Aquatic adaptation and the evolution of smell and taste in whales

Takushi Kishida, J. G. M. Thewissen, Takashi Hayakawa et al. · 9 Feb 2015

INTRODUCTION: While olfaction is one of the most important senses in most terrestrial mammals, it is absent in modern toothed whales (Odontoceti, Cetacea). Furthermore, behavioral evidence suggests that gustation is very limited. In contrast, their aquatic sistergroup, baleen whales (Mysticeti) retain small but functional olfactory organs, and nothing is known about their gustation. It is…

Evolution of the vertebrate skeleton: morphology, embryology, and development

Tatsuya Hirasawa, Shigeru Kuratani · 12 Jan 2015

Two major skeletal systems-the endoskeleton and exoskeleton-are recognized in vertebrate evolution. Here, we propose that these two systems are distinguished primarily by their relative positions, not by differences in embryonic histogenesis or cell lineage of origin. Comparative embryologic analyses have shown that both types of skeleton have changed their mode of histogenesis during evolution. Although…

High diversity in species, reproductive modes and distribution within the Paramacrobiotus richtersi complex (Eutardigrada, Macrobiotidae)

Roberto Guidetti, Michele Cesari, Roberto Bertolani et al. · 3 Jan 2019

For many years, Paramacrobiotus richtersi was reported to consist of populations with different chromosome numbers and reproductive modes. To clarify the relationships among different populations, the type locality of the species (Clare Island, Ireland) and several Italian localities were sampled. Populations were investigated with an integrated approach, using morphological (LM, CLSM, SEM), morphometric, karyological, and…

Planarian shows decision-making behavior in response to multiple stimuli by integrative brain function

Takeshi Inoue, Hajime Hoshino, Taiga Yamashita et al. · 28 Jan 2015

INTRODUCTION: Planarians belong to an evolutionarily early group of organisms that possess a central nervous system including a well-organized brain with a simple architecture but many types of neurons. Planarians display a number of behaviors, such as phototaxis and thermotaxis, in response to external stimuli, and it has been shown that various molecules and neural…

Zoological Letters template — frequently asked questions

How do I write a paper in the Zoological Letters format?
In DocuGuru you write your manuscript in a normal editor — no LaTeX setup required — and select the Zoological Letters template. When you export, DocuGuru compiles the paper into the official BMC (BioMed Central) format and hands you a submission-ready PDF along with the editable LaTeX source.
What reference style does Zoological Letters use?
Zoological Letters uses Numbered (Vancouver) references, shown as numbered [1], [2] markers in the text. DocuGuru formats every in-text citation and the reference list in this exact style automatically. A reference appears like this: 1. Smith A, Jones B, Lee C. A representative article title. Zoological Letters. 2023;12(3):45–58.
Do I need to know LaTeX to submit to Zoological Letters?
No. DocuGuru generates the bmcart LaTeX class and compiles the PDF for you in the background, so you get a BMC (BioMed Central)-ready Zoological Letters document without writing any LaTeX. If you do want it, the LaTeX source is included in the export.
Can I import an existing draft into the Zoological Letters template?
Yes. Paste or upload your current manuscript — Word, LaTeX, Markdown, or plain text — and DocuGuru reflows it into the Zoological Letters format with correct headings, figures, tables, and numbered citations.
What is Zoological Letters?
BMC / BioMed Central journal. It is a life sciences and medicine journal, published by BMC (BioMed Central).
Can I export a submission-ready Zoological Letters PDF?
Yes — DocuGuru produces a PDF built with the official Zoological Letters template (the bmcart class) that is ready to submit to BMC (BioMed Central), together with the matching LaTeX source files.
How much does the Zoological Letters template cost?
You can start writing in the Zoological Letters template for free. Exporting the final submission-ready Zoological Letters PDF and LaTeX source is part of DocuGuru's paid plans — see the app for current pricing.
Use the Zoological Letters template